By David Johns
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Nonetheless, change is also not wholesale, allowing researchers to extrapolate from a mechanism in one organism to the counterparts of that mechanism in its descendants. In some cases the conservation is remarkable. The structure of the genetic regulatory genes that determine segment identity in Drosophila is also observed in humans; and the Pax-6 genes that regulate eye development in the octopus are clearly homologues of the Pax-6 genes that regulate eye development in humans and in Drosophila.
According to this sense, mechanistic explanation ceases to be relevant, since there are no component parts and operations to be distinguished. The organization of the system is then the focus of explanations of systemic behavior. The second sense, which we developed in this section, embraces mechanistic explanation and recognizes the importance of characterizing the component parts and operation of the relevant mechanism, but treats the phenomenon produced by the mechanism as emergent when the operation of the components cannot be predicted from how they operate in simpler systems and the nonlinearities affecting their behavior affects the behavior of the whole system.
Though such systems are fully naturalistic, there is still space for emergence in a sense that is not merely epistemic. Again there are two conditions. If these are met as well as the first two, there will be significant naturalistic emergence. First, we have already noticed that in nonsequential systems, what we called a common assumption of decomposition will fail. The operations of component parts in the system will depend on the actual behavior and the capacities of other components; and to the extent that feedback is systemwide, these dependencies will result in operations that are specific to the system.
A beginner's guide to Forth by David Johns
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